Preprints

Baral, S., Halali, S., Ittonen, M., Fisher, K., Gotthard, K. & Wheat, C. W. A macroevolutionary gene network reveals diapause evolutionary dynamics beyond the circadian clock and predicts microevolution. EcoEvoRxiv. https://doi.org/10.32942/X27664.

Bovolenta, M., Ittonen, M.* & Gotthard, K.* Timing of reproduction modulates fitness effects of local seasonal adaptation in a range expanding butterfly. bioRxiv. https://doi.org/10.64898/2026.01.28.702196.

Peer-reviewed articles

Ittonen, M.* & von Schmalensee, L.* (2026). Rethinking diapause timing thresholds in a warming world. Current Opinion in Insect Science. https://doi.org/10.1016/j.cois.2026.101530.

Ittonen, M.*, Nielsen, M. E.*, Siemers, I., Friberg, M. & Gotthard, K. (2025). Winters restrict a climate change-driven butterfly range expansion despite rapid evolution of seasonal timing traits. Proceedings of the National Academy of Sciences 122: e2418392122. https://doi.org/10.1073/pnas.2418392122.

von Schmalensee, L., Ittonen, M., Brødsgaard Shoshan, A., Roberts, K.T., Siemers, I., Süess, P., Wiklund, C. & Gotthard, K. (2024). Methodological artefacts cause counter-intuitive evolutionary conclusions in a simulation study. Ecology Letters 26: e14439. https://doi.org/10.1111/ele.14439

Ittonen, M., Roberts, K.T., Lehmann, P. & Gotthard, K. (2023). A range-expanding butterfly is susceptible to cold and long winters but shows no signs of local adaptation to winter conditions. Functional Ecology 37: 3064–3078. https://doi.org/10.1111/1365-2435.14445

Ittonen, M., Hagelin, A., Wiklund, C. & Gotthard, K. (2022). Local adaptation to seasonal cues at the fronts of two parallel, climate‐induced butterfly range expansions. Ecology Letters 25: 2022–2033. https://doi.org/10.1111/ele.14085

Hakala, S. M.*, Ittonen, M.*, Seppä, P. & Helanterä, H. (2020). Limited dispersal and an unexpected aggression pattern in a native supercolonial ant. Ecology and Evolution 10: 3671–3685. https://doi.org/10.1002/ece3.6154

* Joint first or last authors